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Chain drive Model A

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It looks to me like it's not for four wheel drive. There appears to be a pivot in the center under the upper beam, so if you wanted to change direction on the rails, you could just turn the entire beam and car body 180 degrees, and drive the other way!

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Mileage might be surprisingly good. In a railroad trade magazine from the eighties I read that a Chev Suburban on steel wheels, got 60-65 MPG in track inspection and maintenance duties. The steel wheels have much less resistance than rubber tires, the grades are gentle, and they don't go very fast.

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